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"Evolution of complex floral trait mediated by generalized pollination networks" Generalist plants interact with constellations of pollinators varying in their frequency of interaction and fitness impact. These heterogeneous groups of organisms are themselves networks that define the pollination niches of plants. The composition, topology and architecture of these pollination networks vary spatially, temporally and across organization levels: within individuals phenologically, among individuals of the same population, among nearby populations, among geographical regions, among related species and macroevolutionarily among phylogenetically-distant species. How generalist plants evolve in response to pollinators in this fractal context is yet an unsolved question. Here, using as model system plant species of the Brassicaceae family, I explore the role played by pollinator networks in the micro- and macroevolution of generalist plants. At microevolutionary level, several attributes of individual-based pollination networks favor the occurrence of significant selection on plant phenotype and promote the maintenance of a fine-scale genetic structure. But not only organisms may be agent of selection. The whole individual network itself can constitute as selective pressures, with some network metrics becoming part of the extended interactive phenotype of generalist plants. Because the inherent spatial variability of individual networks, selection geographic mosaics and landscape-scale local adaptations easily emerge in generalist systems. Scaling up, spatial variation in the attributes of pollinator networks may trigger pollinator-mediated ecological speciation or, on the contrary, foster the occurrence of hybrid zones. At the macroevolutionary level, the lability of clade-oriented networks may cause the occurrence of pollinator-mediated phenotypic diversification. All this evidence suggests that the patterns of plant evolution in generalist scenarios are more complex than previously thought. Their study from a network perspective provides us with a richer and sounder picture of the role that multispecific assemblages of pollinators play in the phenotypic evolution of generalist plants.

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